(pronounced: good-YEAR-ah)


Cranichideae subtribe Goodyerinae

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Succulent trailing terrestrials rooting at the nodes. Leaves in a rosette or scattered along stem, petiolate, often variously colored and variegated. Inflorescences terminal subsessile to long-penculate erect racemes, often pubescent. Flowers globose or tubular, often secund, often barely opening, white, green or reddish-brown. Dorsal sepal and petals appressed forming a hood or banner over the column, the lateral sepals free, sometimes spreading-divergent. Lip unlobed, saccate, with long internal trichomes. Column short, withour wings or foot, the rostellum elongate, deeply bifid; pollinia 2, often deeply cleft, on a common elongate viscidium.


Honoring the early English botanists John Goodyer.


A cosmopolitan genus of about 55 species, absent in Africa. Asian species of Goodyera with variegated leaves are a minor component of traditional Chinese medicine.

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Ackerman, J. D. 1975. Reproductive biology of Goodyera oblongifolia (Orchidaceae). Madroño 23(4):191-198.

Alexander, C.and G. Hadley 1984. The effect of mycorrhizal infection of Goodyera repens and its control by fungicide. New Phytol. 97:391-400.

Alexander, C. and G. Hadley 1985. Carbon movement between host and mycorrhizal endophyte during the development of the orchid Goodyera repens Br. New Phytol. 101:657-665.

Alexander, C., I. J. Alexander and G. Hadley 1984. Phosphate uptake by Goodyera repens in relation to mycorrhizal infection. New Phytol. 97:401-411.

Champlin, R. L. 1976. Scotch pine as an associate of the tesselated rattlesnake plantain. Rhodora 78:788-789.

Fernald, M. L. 1899. The rattlesnake plantains of New England. Rhodora 1:2-7.

Healy, P. ###et al. 1980. Morphology of orchid seeds, III. Native California and related species of Goodyera, Piperia, Platanthera, and Spiranthes. Amer. J. Bot. 67(4):508-518.

Kallunki, J. A. 1976. Population studies in Goodyera (Orchidaceae) with emphasis on the hybrid origin of G. tesselata. Brittonia 28(1):53-75.

Kallunki, J. 1981. Reproductive biology of mixed-species populations of Goodyera (Orchidaceae) in northern Michigan. Brittonia 33(2):137-155.

Purves, S. and G. Hadley 1976. The physiology of symbiosis in Goodyera repens. New Phytol. 77:689-696.

Sharma, M. and S. P. Vij 1984. Embryological studies in Orchidaceae. 3. Goodyera repens R. Br. Res. Bull. Panjab Univ., n. s., Sci. 35(1-2):61-64.

Sood, S. K. 1988. Development of gametophytes, embryogeny and pericarp in Goodyera repens (Orchidaceae, Neottieae). Proc. Indian Acad. Sci., Plant Sci. 98:149-156.

Wong, K. C. and M. Sun 1999. Reproduction biology and conservation genetics of Goodyera procera (Orchidaceae). Amer. J. Bot. 86:1406-1413.

Xu, S.-X., Y.-H. Xiao and Z.-D. Yang 1987. Studies on the ontogeny of the pollinium of Goodyera procera. Acta Bot. Sinica 29:577-579.